I am professor in the Department of Zoology, where I carry research out research on a wide variety of topics, from population genetic theory (models of selection and the maintenance of genetic variation; models of selection on non-standard inheritance systems; models of phenotypic plasticity), to biogeography (long-distance dispersal in marine invertebrates) and phylogeny (of cormorants and molluscs), to the history of eugenics. My obtained my PhD from Harvard University, studying under Richard Lewontin; my undergraduate degree was in mathematics from the University of Auckland, from which I also graduated with an MSc in zoology, studying with David Lambert and Brian McArdle.