PhD in population genetics (evolution of sex). Edinburgh. Fellowship 1: in bacteria-phage coevolution. UCSD. 7 years as Editor of TREE Fellowship 2: Evolutionary epidemiology of HIV (within and between host, and linking scales). Modelling. Imperial Fellowship 3: and Group Leader position. Evolutionary epidemiology of human chronic viruses (HIV, HCV, HBV). Modelling, evolutionary analysis of sequencing data, assimilating global HLA data. Oxford.